Squeeze handfuls of spinach to remove as much liquid as possible, then finely chop.
Cook onion and garlic in butter in a large heavy skillet over moderate heat, stirring, until soft, 4 to 5 minutes. Add mushrooms, 1/2 teaspoon salt, 1/4 teaspoon pepper, and nutmeg and cook, stirring, 5 minutes. Stir in spinach, then remove from heat.
Spread one third of bread cubes in a buttered 3-quart gratin dish or other shallow ceramic baking dish and top evenly with one third of spinach mixture. Sprinkle with one third of each cheese. Repeat layering twice (ending with cheeses).
Whisk together milk, eggs, mustard, and remaining 1/2 teaspoon salt and 1/4 teaspoon pepper in a large bowl and pour evenly over strata. Chill strata, covered with plastic wrap, at least 8 hours (for bread to absorb custard).
Preheat oven to 350°F. Let strata stand at room temperature 30 minutes.
Bake strata, uncovered, in middle of oven until puffed, golden brown, and cooked through, 45 to 55 minutes. Let stand 5 minutes before serving.
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